Hoxb5 marks long-term haematopoietic stem cells and reveals a homogenous perivascular niche.

Publication Type Academic Article
Authors Chen J, Miyanishi M, Wang S, Yamazaki S, Sinha R, Kao K, Seita J, Sahoo D, Nakauchi H, Weissman I
Journal Nature
Volume 530
Issue 7589
Pagination 223-7
Date Published 02/11/2016
ISSN 1476-4687
Keywords Hematopoietic Stem Cells, Homeodomain Proteins, Stem Cell Niche
Abstract Haematopoietic stem cells (HSCs) are arguably the most extensively characterized tissue stem cells. Since the identification of HSCs by prospective isolation, complex multi-parameter flow cytometric isolation of phenotypic subsets has facilitated studies on many aspects of HSC biology, including self-renewal, differentiation, ageing, niche, and diversity. Here we demonstrate by unbiased multi-step screening, identification of a single gene, homeobox B5 (Hoxb5, also known as Hox-2.1), with expression in the bone marrow that is limited to long-term (LT)-HSCs in mice. Using a mouse single-colour tri-mCherry reporter driven by endogenous Hoxb5 regulation, we show that only the Hoxb5(+) HSCs exhibit long-term reconstitution capacity after transplantation in primary transplant recipients and, notably, in secondary recipients. Only 7-35% of various previously defined immunophenotypic HSCs are LT-HSCs. Finally, by in situ imaging of mouse bone marrow, we show that >94% of LT-HSCs (Hoxb5(+)) are directly attached to VE-cadherin(+) cells, implicating the perivascular space as a near-homogenous location of LT-HSCs.
DOI 10.1038/nature16943
PubMed ID 26863982
PubMed Central ID PMC4854608
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