Translaminar synchronous neuronal activity is required for columnar synaptic strengthening in the mouse neocortex.

Publication Type Academic Article
Authors Vargas-Ortiz J, Lin L, Martinez V, Liu R, Babij R, Duan Z, Wacks S, Sun L, Wang A, Khan S, Soto-Vargas J, De Marco García N, Che A
Journal Nat Commun
Volume 16
Issue 1
Pagination 1296
Date Published 02/03/2025
ISSN 2041-1723
Keywords Neocortex, Somatosensory Cortex, Pyramidal Cells, Synapses, Neurons
Abstract Synchronous neuronal activity is a hallmark of the developing mouse primary somatosensory cortex. While the patterns of synchronous neuronal activity in cortical layer 2/3 have been well described, the source of the robust layer 2/3 activity is still unknown. Using a novel microprism preparation and in vivo 2-photon imaging in neonatal mice, we show that synchronous neuronal activity is organized in barrel columns across layers. Monosynaptic rabies tracing and slice electrophysiology experiments reveal that layer 2/3 pyramidal neurons receive significant layer 5 inputs during the first postnatal week, and silencing layer 5 synaptic outputs results in a significant reduction in spontaneous activity, abnormal sensory-evoked activity and disrupted layer 4-layer 2/3 connectivity. Our results demonstrate that translaminar layer 5-layer 2/3 connectivity plays an important role in synchronizing the developing barrel column to ensure the strengthening of layer 4-layer 2/3 connections, supporting the formation of the canonical cortical organization in barrel cortex.
DOI 10.1038/s41467-024-55783-w
PubMed ID 39900899
PubMed Central ID PMC11791040
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